1MOE Key Laboratory for Membraneless Organelles & Cellular Dynamics, School of Life Sciences, University of Science and Technology of China, Hefei 230026, China 2Anhui Key Laboratory for Chemical Biology and New Quality Medicine, Hefei National Research Center for Physical Sciences at the Microscale, Hefei 230027, China *Correspondence to:Liangyu Zhang , Email:lyzhang0@mail.ustc.edu.cn Xuebiao Yao , Email:yaoxb@ustc.edu.cn